What kind of animal is the pancake turtle (Pancake Tortoise)? An updated explanation of its characteristics, personality, ecology, and habitat.

動物図鑑

What kind of animal is a pancake tortoise? We will explain the characteristics, personality, ecology, habitat, and sales of small animals of amphibious reptiles. This tortoise lives in eastern Africa and is a member of the turtle family that is popular as a pet, but it is actually designated as an endangered species. It is suggested that it may become extinct in the future.

What is Pancake Turtle? Basic Stats

The pancake turtle is a turtle classified in the order Testudins, family Testudinidae, genus Malacochersus. It is a small turtle with a carapace length of only about 15 cm. Its scientific name is Malacochersus tornieri, and its English name is African pancake tortoise. There is a lot of information on the Internet, and it is also sold in stores and is updated regularly.

Japanese(和名)パンケーキガメ
English(英名)African pancake tortoise
scientific name(学名)Malacochersus tornieri
classification(分類)Reptilia、 Testudines、 TestudinidaeMalacochersus  
爬虫綱、カメ目、リクガメ科、パンケーキガメ属
IUCN Status(保全状況)CRITICALLY ENDANGERED
Length(体長)15~18cm
Weight(体重)0.5~1kg 

Taxonomy of the Pancake Tortoise (Malaclemys terrapin)

分類階級学名・内容
界(Kingdom)Animalia(動物界)
門(Phylum)Chordata(脊索動物門)
綱(Class)Reptilia(爬虫綱)
目(Order)Testudines(カメ目)
科(Family)Testudinidae(リクガメ科) *一部では海岸ガメに近い分類もあり
属(Genus)Malaclemys(マラクレミス属)
種(Species)Malaclemys terrapin

What are the roots and origins of the pancake tortoise?

🌍 Place of Origin

Its natural distribution is primarily in:

🇹🇿 Tanzania

🇰🇪 Kenya

🇿🇲 Northern Zambia

A particularly distinctive feature is its use of rocky areas, crevices, and spaces beneath rocks, rather than just the open ground typical of most tortoises. Rocky outcrops (kopjes) scattered across the dry savannas and shrublands of East Africa serve as vital habitats.

🧬 The Biggest Mystery: Why Is It So Flat?

This is the most fascinating aspect of the pancake tortoise’s evolution.

Typical tortoises
→ Hard, dome-shaped shell

Pancake tortoise
→ Extremely thin, flexible shell

They possess forms that are nearly polar opposites.

Current research suggests that the pancake tortoise evolved from a lineage sharing a common ancestor with the Indotestudo and Testudo genera. However, no fossils representing the pancake tortoise’s direct ancestors have been discovered to date, leaving a significant gap in its evolutionary history.

An Evolutionary Scenario

Typical ancestral tortoise

Competition and predation pressure

Individuals utilizing rocky areas gain an advantage

Escaping into rock crevices and under rocks

Body shapes that fit into narrower gaps become advantageous

Shell becomes flatter

Shell ossification decreases, increasing flexibility

🐢 Modern pancake tortoise

This evolutionary scenario is a leading hypothesis. Researchers propose that the shift toward utilizing rock crevices—driven by predation pressure—likely led to the development of the species’ current, extremely flat body shape.

🪨 A “Flat Shell”: Not a Weakness, but a Weapon

For the pancake tortoise, its flat body is considered an adaptation to a specialized life among rocks.

Squeezing into a rock crevice

Pushing the body deep into the gap

Wedging itself in place by expanding its body

These behaviors are made possible by its unique anatomy. Consequently, unlike typical tortoises that possess hard, rounded shells, this species has evolved to specialize in using rock crevices themselves as refuges.

🔬 More Fascinating Recent Research

A 2026 study of mitochondrial genomes revealed a significant genetic divergence between the Kenyan and Tanzanian lineages, suggesting they may have diverged approximately 5.74 million years ago.

Researchers suggest this could serve as a basis for considering the existence of “cryptic species” in the future; however, they are not currently formally classified as separate species.

Habitat

Pancake turtles are found in parts of Africa, including Kenya, Zambia, and Tanzania.

  1. Geographic Distribution

Distributed primarily in estuaries, brackish waters, and salt marshes along the east coast of North America.

State Distribution Example:

Coastal areas from Maine to Florida

Prefers brackish waters, tidal flats, and salt marshes along the coast.

  1. Habitat Type

Mainly lives in brackish waters (estuaries and tidal flats where freshwater and seawater mix).

Often sunbathes on sand or mud.

Feeds, rests, and spawns in shallow waters or areas with thick saltweed.

  1. Relationship with Water

Semi-aquatic, primarily active in the water.

Excellent swimmers, primarily foraging and escape behaviors in the water.

Comes onto land to sunbathe, but quickly dives back into the water to escape.

  1. Habitat Characteristics

Adapted to specialized environments such as coastal tidal flats and salt marshes.

Often hides in shallow waters, aquatic plants, sand, and mud to escape predators.

Prefers coastal areas with stable air and water temperatures.

What are its characteristics? What kind of creature is it?

The pancake turtle’s shell is flat, thin and soft like a pancake. That’s why it was named pancake. The flat and elastic shell makes it easier for it to hide in narrow gaps. Its limbs are sturdy and its claws are sharp and strong. The color of its dorsal shell is yellow or yellowish brown, and the color of its plastron is yellowish brown. Each shell has dark spots with radial spots around them. Pancake turtles live in savannas, forests and brush from lowlands to mountainous areas.

  1. Body Size and Morphology

Body Length: Approximately 15-23 cm (medium-sized turtle) in adults

Weight: Approximately 0.5-1.5 kg

Shell (Carapace):

Very flat and soft (allows for easy escape and quick movement in the water)

Carapace color is dark brown to blackish brown, pattern varies from individual to individual

Head and Limbs:

Small head, large eyes, and well-developed senses

Webbed limbs for swimming and underwater movement

  1. Behavior and Personality

Semi-aquatic and excellent swimmers

Timid and cautious, they will immediately retreat into the water if they sense danger

Rarely aggressive, with a gentle personality

  1. Diet

Omnivorous

Food:

Small animals such as shellfish, crustaceans, small fish, and insects

Also eats plant foods such as aquatic plants, algae, and fruit

Feeding Method:

Extends its neck to search for food in the water and chews it with its jaws

  1. Biological Impressions

A semi-aquatic turtle characterized by its quick movements and escape ability in the water

With its flat shell and small body, it can quickly move through shallow waters and tidal flats

Timid and cautious, its behavior is primarily to flee from humans and predators

What is its personality like?

Pancake turtles are quite cautious turtles. They often hibernate in hiding places such as under rocks or in rock crevices, and when they sense danger, they quickly escape to their hiding places. They never stray far from their homes and are active in the early morning and at dusk.

  1. Timid and cautious

If they sense danger, they will immediately retreat into the water.

Extremely wary of humans and predators.

Even when on land, they will dive into the water at the slightest provocation.

  1. Gentle and low-aggression

Very rarely aggressive, they rarely bite.

Tends to be docile and quiet.

  1. Low sociality

Generally solitary.

Only encounters other individuals at feeding sites or sunbathing areas.

Rarely lives in groups.

  1. Adaptive behavior

Excellent at quickly escaping from shallow waters and tidal flats.

Excellent at moving around and using hiding places underwater.

What is their ecology like?

Pancake turtles are herbivorous and live by eating grass and other things. They breed oviparously, with males fighting over females. They mate year-round except during the dry season. They lay one egg at a time. The incubation period is 4-7 months, and they live for about 30 years.

  1. Lifestyle

Semi-aquatic, primarily living underwater

Mainly active in brackish waters, estuaries, tidal flats, and salt marshes

Coming onto land only for basking and spawning

Feeding and moving around by swimming underwater

  1. Diet

Omnivorous

Food:

Small animals such as shellfish, crustaceans, small fish, and insects

Also consume plant foods such as aquatic plants, algae, and fruit

Feeding Method:

Extends its neck to catch food from the water and crushes it with its jaws

Adapted to a semi-aquatic environment, it can efficiently search for food underwater

  1. Reproduction and Rearing

Breeding Season: Mainly spring to early summer (varies by region)

Oviparous. Females come onto sandy beaches or tidal flats, dig burrows, and lay eggs.

Egg count: Approximately 5-15 eggs per clutch (varies by individual)

Incubation period: Approximately Around 60 days

Juvenile Independence:

They survive independently in the water immediately after hatching.

They grow up in a safe place, being wary of predators.

  1. Behavior and Sociality

Primarily solitary.

Interaction with other individuals is limited to sunbathing areas and breeding season.

Their range of movement is shallow waters and tidal flats, varying depending on water levels and food distribution.

  1. Habitat Relationship

Adapted to estuaries, tidal flats, and salt marshes along the east coast of North America.

They primarily feed, move, and use hiding places in the water.

They come onto land only for spawning and sunbathing.

Do they have any natural enemies?

Pancake turtles are small turtles, so most carnivores are natural enemies.

About Pancake Turtle Hatches

This article provides a detailed overview of the characteristics and ecology of pancake turtle (Malaclemys terrapin) hatchlings (from hatching to juvenile stage).

  1. Birth and Hatching

Oviparous: Females dig burrows in sandy beaches or tidal flats and lay eggs there.

Number of eggs: Approximately 5-15 per clutch (varies by individual).

Incubation Period: Approximately 50-60 days (depending on temperature and humidity).

Body Length and Weight:

Body Length: Approximately 3-5 cm

Weight: Approximately 20-50 g

Appearance:

Carapace is soft and flat.

Dark brown to blackish brown in color, with pale markings.

Webbed feet and swimming ability are apparent immediately after hatching.

  1. Juvenile Behavior

Highly independent: Lives in the water immediately after hatching

Shy and cautious:

Quickly retreats into the water when it senses a predator or human presence

Protects by hiding in the shade of aquatic plants, sandy bottoms, or tidal flats

Feeding Behavior:

Eat small shellfish, insects, and aquatic plants on their own

Extends their neck to search for food in the water and crushes it with their jaws

  1. Growth Stage

Immediately after hatching

Solitary, prioritizes hiding

Juvenile Stage (several months to 1 year)

Their carapace hardens, and their patterns and colors gradually become more vivid

Increased feeding and range of activity

Juvenile Stage (1-2 years)

Improved underwater swimming and escape ability

Body length and weight increase as they approach adulthood

  1. Habitat Relationship

They use safe areas such as shallow waters, tidal flats, and the shade of aquatic plants or sandy bottoms.

Protection from predators is paramount.

They primarily spend their time underwater and rarely come onto land.

Is the pancake turtle an endangered species?

The pancake turtle is listed in Appendix I of the Convention on International Trade in Endangered Species of Wild Fauna and Flora, and international trade is restricted. Not only is its habitat decreasing due to recent development, but it is also overfished as a pet, which is a problem. If you want to keep it as a pet, it will cost you money as its price has risen sharply.

  1. IUCN Assessment

Classified as Near Threatened (NT) on the IUCN Red List.

While populations remain relatively stable in some areas, habitat destruction and human activities pose a risk of decline.

  1. Causes of Decline

Habitat Destruction

Estuarine and Tidal Flat Landfills, Revetment Construction, and Urban Development

Traffic and Human Interference

Vehicles and Pedestrians Endanger Basking Individuals

Changes in Predation Pressure

Impact of Predators such as Cats, Dogs, and Birds

Climate Change

Flooding of Nesting Holes Due to Low and High Tides and Storms Reduces Hatching Rates

  1. Conservation Efforts

Establishment of Protected Areas: Preservation of Tidal Flats and Salt Marshes

Breeding Site Management: Protection of Nesting Holes During the Spawning Season

Education and Awareness-Raising Activities: Raising Conservation Awareness among Local Residents

Can you keep a pancake turtle as a pet?

It is possible to keep a pancake turtle as a pet. The following items are necessary for keeping a turtle. Small individuals are newly registered and sold, and there are many adoption requests, so check them out. Pay attention to the type of food and cage. They also have a long lifespan, so take that into consideration. Check the food and how to keep them warm. There are many products available in shops, and it is easy to purchase supplies such as food, cases, and lights.

Cage

Depending on the size of the shell and the growth rate, a cage with a width of about 60 cm is required at the shop. It is recommended to use a reptile cage made of glass, acrylic, or plastic.

Flooring

There are various types of flooring, from large to small. There are many types such as red clay, coconut shells, and artificial turf. They are sometimes sold at low prices during campaigns.

Heater

If necessary, prepare an air conditioner or heater to make sure your turtle is comfortable. There are panel heaters, spot lamps, and basking lights.

Thermo-hygrometer

When raising amphibious turtles, it is very important to control the temperature and humidity. It is best to keep the temperature at around 30 degrees.

Lighting

Turtles are animals that maintain their health by exposing themselves to strong ultraviolet rays throughout the day, producing vitamin D3 in their bodies that promotes calcium absorption. For this reason, lighting is essential. There are incandescent and fluorescent tube types, and it is best to choose one that suits the cage. Use the sunlight hours as a guide for lighting.

Shelter

Turtles are living creatures, so it is a good idea to have a shelter as a point to create a place for them to escape indoors. Make sure to install it properly so as not to cause them too much stress.

Water dish

A water dish is also essential. Clean the water bowl every day and provide fresh water.

Pet food

Turtles are herbivorous and their food consists mainly of small vegetables and fruits, with some komatsuna and mushrooms. Some turtles eat wild plants instead of vegetables. Give them fruits such as bananas, watermelons, apples, strawberries, kiwis, and mangoes as snacks. Some people also give supplements as a reference. It is a good idea to start by offering commercially available products. These contain all the nutrients that living creatures need.

Zoos Housing the Species (Japan & Worldwide)

🇯🇵 Japan

Kyoto City Zoo (Kyoto Prefecture) ✅ Confirmed
Toba Aquarium (Mie Prefecture) ✅ Records available
Tennoji Zoo (Osaka Prefecture) ✅ Confirmed

Kyoto City Zoo lists the Pancake Tortoise (Malacochersus tornieri) in its official animal encyclopedia.

Toba Aquarium is also listed as a facility housing the species in global zoo databases.

Regarding Tennoji Zoo, the housing of two Pancake Tortoises can be confirmed in their official annual report (FY2022 data).

Additionally, Pancake Tortoises are listed as an “International Rare Species of Wild Fauna and Flora” in documents from the Ministry of the Environment regarding certified facilities for the conservation of rare species.

🌎 Worldwide

The species is kept in a significant number of zoos around the world.

🇺🇸 USA

Zoo Atlanta

Bronx Zoo

San Diego Zoo

San Diego Zoo Safari Park

Detroit Zoo

Cincinnati Zoo & Botanical Garden

St. Louis Zoo

Oklahoma City Zoo

Tulsa Zoo

Memphis Zoo

Dallas Zoo

Fort Worth Zoo

Buffalo Zoo

Montgomery Zoo

Tennessee Aquarium

Phoenix Zoo

Oakland Zoo

Birmingham Zoo

National Zoological Park (Smithsonian’s National Zoo)

Records of Pancake Tortoises being kept at numerous facilities in the US can be confirmed via Zooinstitutes.

🇩🇪 Germany

Aquazoo Löbbecke Museum Düsseldorf

Zoo Frankfurt

Zoo Leipzig

Zoo Berlin

Tierpark Berlin

Zoo Wuppertal

Zoo Hannover

Zoo Magdeburg

Zoopark Erfurt

Tierpark Hellabrunn (Munich)

Zoo Schwerin

Germany has a particularly large number of facilities housing the species; many can also be confirmed via Zootierliste. 🇳🇱 Netherlands

Rotterdam Zoo (Diergaarde Blijdorp)

ARTIS Amsterdam

🇧🇪 Belgium

ZOO Antwerpen

Pairi Daiza

🇬🇧 UK

West Midland Safari Park

Chester Zoo

Dudley Zoo

Dartmoor Zoological Park

Cotswold Wildlife Park

Twycross Zoo

Linton Zoo

Belfast Zoo

🇫🇷 France

ZooParc de Beauval

🇨🇭 Switzerland

Zoo Zürich

Zoo Basel

🇨🇿 Czech Republic

Prague Zoo

Zoo Plzeň

🇵🇱 Poland

Poznań Old Zoo

Wrocław Zoo

🇫🇮 Finland

Helsinki Zoo

🇮🇹 Italy

Parco Natura Viva

🇭🇺 Hungary

Budapest Zoo

🇷🇺 Russia

Moscow Zoo

Perm Zoo

Roev Ruchey Zoo

Yaroslavl Zoo

🇨🇳 China

Beijing Zoo

🇺🇦 Ukraine

Kyiv Zoo

EKZOland

🇰🇿 Kazakhstan

Almaty Zoo

🇹🇭 Thailand

Monsters Aquarium Pattaya

🇦🇪 UAE

Sharjah Safari

These are facilities that can be verified in global databases of zoological facilities.

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